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Title: A Study of Stomatal Structure in Pandanaceae 
Author: Tomlinson, P.B.
Date: 1965-01
Publisher: University of Hawai'i Press
Citation: Tomlinson PB. 1965. A study of stomatal structure in Pandanaceae. Pac Sci 19(1): 38-54.
Abstract: Many species of Pandanaceae have unspecialized stomata. In species
of Pandanus there is a range of stomatal structure involving increasing elaboration
of papillae arising from subsidiary and neighbouring cells so that guard cells, which
are otherwise constant in structure throughout the family, are increasingly provided
with a canopy of papillae which, together with sinking of the stomatal apparatus,
produces an outer stomatal chamber. In the most specialized stomata this chamber
is itself virtually occluded by lobed papillae.
Freycinetia may show significant differences from Pandanus. This linear trend
of specialization can often be partly or wholly demonstrated on a single individual
because stomata are always unspecialized on the first scalelike leaves of each renewal
shoot, thereafter being increasingly specialized on subsequent leaves until the maximum
potential elaboration characteristic of the species is realised on foliage leaves.
This is briefly discussed in relation to taxonomy, physiology, and ecology.
ISSN: 0030-8870

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