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Title: Factors Affecting the Distribution of Tylos punctatus (Isopoda, Oniscoidea) on Beaches in Southern California and Northern Mexico 
Author: Hayes, Willis B.
Date: 1977-04
Publisher: University of Hawaii Press
Citation: Hayes WB. 1977. Factors affecting the distribution of Tylos punctatus (Isopoda, Oniscoidea) on beaches in southern California and northern Mexico. Pac Sci 31(2): 165-186.
Abstract: Populations of the isopod Tylos punctatus were studied at five
widely separated beaches between Los Angeles, California, and Punta Banda,
Baja California (Mexico). Population densities at these sites were 10 3-10 5
animals per meter of beach frontage; smaller populations were not located,
although more than 30 intermediate sites were sampled.
Cross-beach distribution appears to be related to sand moisture content
rather than sand coarseness. Longshore distribution on a given beach is patchy,
but the causative factors for this were not determined.
Studies of the isopods' population dynamics on two beaches indicated that
the populations were increasing, with doubling times of 2-12 years. If these
are minimum values, it is hypothesized that the discontinuous distribution of
the animals on this coast can represent a balance between catastrophic beach
erosion and the natural ability of the populations to regenerate and disperse.
ISSN: 0030-8870

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